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Professor John Maynard Smith,[1] F.R.S. (6 January 1920 – 19 April 2004) was a British theoretical evolutionary biologist and geneticist. Originally an aeronautical engineer during the Second World War, he then took a second degree in genetics under the well-known biologist J.B.S. Haldane. Maynard Smith was instrumental in the application of game theory to evolution and theorized on other problems such as the evolution of sex and signalling theory. John Maynard Smith was born in London, the son of a surgeon, but following his father's death in 1928 the family moved to Exmoor, where he became interested in natural history. Quite unhappy with the lack of formal science education at Eton College, Maynard Smith took it upon himself to develop an interest in Darwinian evolutionary theory and mathematics, after having read the work of old Etonian J.B.S. Haldane, whose books were in the school's library despite the bad reputation Haldane had at Eton for his communism. On leaving school, Maynard Smith joined the Communist Party of Great Britain and started studying engineering at Trinity College Cambridge. When the second world war broke out in 1939, he defied his party's line and volunteered for service. He was rejected, however, because of poor eyesight and was told to finish his engineering degree, which he did in 1941. He later quipped that "under the circumstances, my poor eyesight was a selective advantage — it stopped me getting shot". The year of his graduation, he married Sheila Matthew, and they were later to have two sons and one daughter (Tony, Carol, and Julian). Between 1942 and 1947 he applied his degree to military aircraft design. Maynard Smith then took a change of career, entering University College London (UCL) to study fruit fly genetics under Haldane. After graduating he became a lecturer in Zoology at UCL between 1952 and 1965, where he directed the Drosophila lab and conducted research on population genetics. He published a popular Penguin book, The Theory of Evolution, in 1958 (with subsequent editions in 1966, 1975, 1993).
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